Seems like the main difference is that you’re counting up with status and counting down with genetic fitness.
There are plenty of people who plan their lives around having & raising kids. And there are plenty of people who put tons of effort into leveling up their videogame characters, or affiliating-at-a-distance with Taylor Swift or LeBron James, with minimal real-world benefit to themselves.
And it’s easier to count up lots of things as status-related if you’re using a vague concept of status which can encompass all sorts of status-related behaviors, including (e.g.) both status-seeking and status-affiliation. “Inclusive genetic fitness” is a nice precise concept so it can be clear when individuals fail to aim for it even when acting on adaptations that are directly involved in reproduction & raising offspring.
I read this as saying ‘a common view is that being criticized is bad because it hurts your reputation, but as a person with some knowledge of the secret lore of rationality I believe that being criticized is good because you can learn from it.’
And he isn’t making a claim about to what extent the existing LW/rationality community shares his view.