A single developmentally modular process can give rise to multiple architectural modules.
To the extend that a single developmental modular process leads to multiple architectual modules, it’s a reasonable argument that it’s improper to analyse the evolutionary advantages of a single architectual module without knowing looking at the other achitectual modules that spring out of the same developmental modular process.
To the extend that a single developmental modular process leads to multiple architectual modules, it’s a reasonable argument that it’s improper to analyse the evolutionary advantages of a single architectual module without knowing looking at the other achitectual modules that spring out of the same developmental modular process.
That sounds like a much more reasonable criticism than the ones presented in the OP.