Do you mean to say that the expected inclusive fitness of a randomly selected creature from the population goes up with time? Well, if we sum that up over the whole population, we obtain the total number of offspring—right? And dividing that by the current population, we see that the expected inclusive fitness of a randomly selected creature is simply the population’s growth rate. The problem is that evolution does not always lead to >1 population growth rate. Eliezer gave a nice example of that: “It’s quite possible to have a new wolf that expends 10% more energy per day to be 20% better at hunting, and in this case the sustainable wolf population will decrease as new wolves replace old.”
Inclusive reproductive fitness.
Do you mean to say that the expected inclusive fitness of a randomly selected creature from the population goes up with time? Well, if we sum that up over the whole population, we obtain the total number of offspring—right? And dividing that by the current population, we see that the expected inclusive fitness of a randomly selected creature is simply the population’s growth rate. The problem is that evolution does not always lead to >1 population growth rate. Eliezer gave a nice example of that: “It’s quite possible to have a new wolf that expends 10% more energy per day to be 20% better at hunting, and in this case the sustainable wolf population will decrease as new wolves replace old.”